The first invention of mankind was the wheel. The invention of the wheel is probably the most important invention of all time.

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问题     The first invention of mankind was the wheel. The invention
of the wheel is probably the most important invention of all time.
Without the wheel, the world simply wouldn’t exist as we know. 【M1】______
The invention of the wheel was at the root of the Industrial
Revolution, which it would take a long time to get there. 【M2】______
    Although no wheel forms are found in nature; undoubtedly the
earliest "wheels" were smooth logs which were used for moving
weights over the earth’s surface. It is a mystery so as to who 【M3】______
invented the first wheel. No one recorded who was or when it 【M4】______
happened; but when the first inventor has placed a wheel on an axle,
mankind began to roll on one place to another. This type of wheel 【M5】______
have been found among Egyptian relics dating back due to 2,000 B.C, 【M6】______
and earlier Chinese civilizations are credited with independent
invention of the same mechanism.
    The wheel so fascinated man that he has spent centuries
building machines around them; yet in over 4,000 years he has not 【M7】______
changed its basic design. All about us we see the spinning shafts,
gears, flywheels, and rotors are the descendants of the first wheel. 【M8】______
The roaring propeller of an aircraft engine, the whirling wheel of a
giant steam turbine, and the hairspring of a tiny watch are
examples of the motorized motion which characterizes our 【M9】______
mechanism world. It is hard to conceive of continuous motion 【M10】______
without the wheel.
【M4】

选项

答案who∧一he

解析 句法错误。从句子结构看,who was应该是一个宾语从句,很显然这个从句缺少一个成分。
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