Our ape-men forefathers had no obvious natural weapons in the struggle for survival in the open. They had neither the powerful t

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问题     Our ape-men forefathers had no obvious natural weapons in the struggle for survival in the open. They had neither the powerful teeth nor the strong claws of the big cats. They could not【C1】______with the bear, whose strength, speed and claws【C2】______an impressive ""small fire"" weaponry. They could not even defend themselves【C3】______running swiftly like the horses, zebras or small animals. If the ape-men had attempted to compete on those terms in the open, they would have been【C4】______to failure and; extinction. But they were【C5】______with enormous concealed advantages of a kind not possessed by any of their competitors.
    In the search【C6】______the pickings of the forest, the ape-men had【C7】______efficient stereoscopic vision and a sense of colour that the animals of the grasslands did not possess. The ability to see clearly at close range permitted the ape-men to study practical problems in a way that lay far【C8】______the reach of the original inhabitants of the grassland. Good long distance sight was【C9】______another matter. Lack of long-distance vision had not been a problem for forest-dwelling apes and monkeys because the higher the viewpoint, the greater the range of sight so【C10】______they had had to do was climb a tree. Out in the open, however, this simple solution was not【C11】______Climbing a hill would have helped, but in many places the ground was flat. The ape-men【C12】______the only possible solution. They reared up as high as possible on their hind limbs and began to walk upright.
    This vital change of physical position brought about considerable disadvantages. It was extremely unstable and it meant that the already slow ape-men became slower still.【C13】______, they persevered and their bone structure gradually became【C14】______to the new, unstable position that【C15】______them the name Homo erectus, upright man."
【C13】

选项 A、However
B、Therefore
C、Meanwhile
D、Subsequently

答案A

解析 联系上下文语境,该空所在句与前一句构成转折关系,从意义上分析,however符合,且其后常用逗号与句子其他成分隔开。因此,答案为A。
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