In the past few decades, remarkable findings have been made in ethology, the study of animal social behavior. Earlier scientists

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问题     In the past few decades, remarkable findings have been made in ethology, the study of animal social behavior. Earlier scientists had【C1】______ that nonhuman social life was almost totally instinctive or fixed by genetics. Much more careful observation has shown that【C2】______ variation occurs among the social ties of most species, showing that learning is a part of social life. That is, the statuses are not solely fixed by the genes.【C3】______ , the learning that occurs is often at an early age in a process that is called imprinting. Imprinting is clearly not instinctive, but it is not quite like the learning of humans; it is something in between the two. An illustration best【C4】______ the nature of imprinting. Once, biologists thought that ducklings followed the mother duck because of instincts. Now we know that, shortly after they hatch, ducklings fix【C5】______ any object about the size of a duck and will henceforth follow it. So ducklings may follow a basketball or a briefcase if these are【C6】______ for the mother duck at the time when imprinting occurs. Thus, social ties can be considerably【C7】______ , even ones that have a considerable base【C8】______ by genetics.
    Even among the social insects something like imprinting must influence social behavior. For example, biologists once thought bees communicated with others purely【C9】______ instinct. But, in examining a "dance" that bees do to indicate the distance and direction of a pollen source, observers found that bees raised in isolation could not communicate effectively. At a higher level, the genetic base seems to be much more for an all-purpose learning rather than the more specific responses of imprinting. Chimpanzees, for instance, generally【C10】______ very good mother but Jane Goodall reports that some chimps carry the infant upside down or otherwise fail to nurture the young.
【C3】

选项 A、What’s more
B、Hence
C、But
D、However

答案D

解析 A项What’s more可表示递进;B项Hence意为“因此,所以”,表示因果关系。文中上一段提到“学习是动物的社会生活的一部分”,第二段接着讲“学习发生在动物小时候的印象中期”,两段间不存在递进或因果关系。But在表转折时一般不与句子分开,所以D项However为正确答案。
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