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  3. In the past few decades，remarkable findings have been made in ethology，the study of animal social behavior．Earlier scientists ha

In the past few decades，remarkable findings have been made in ethology，the study of animal social behavior．Earlier scientists ha

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问题   In the past few decades，remarkable findings have been made in ethology，the study of animal social behavior．Earlier scientists had【1】that nonhuman social life was almost totally instinctive or fixed by  genetics．Much more careful observation has shown that【2】variation occurs among the social ties of most species，showing that learning is a part of social life．That is，the【3】are not solely fixed by the genes．
  【4】，the learning that occurs is often at an early age in a process that is called imprinting．Imprinting is clearly【5】instinctive，but it is not quite like the learning of humans；it is something in between the two．An illustration best【6】the nature of imprinting．Once，biologists thought that ducklings followed the mother duck because of instincts．Now we know that，shortly【7】they hatch．ducklings fix【8】any object about the size of a duck and will henceforth follow it．So ducklings may follow a basketball or a briefcase if these are【9】for the mother duck at the time when imprinting occurs．Thus，social ties can be considerably【10】，even ones that have a considerable base 【11】 by genetics．
  Even among the social insects something like imprinting 【12】influence social behavior．For example，biologists once thought bees communicated with others purely 【13】instinct．But，in examining a“dance”that bees do to indicate the distance and direction of a pollen source，observers found that bees raised in isolation could not communicate effectively．At a higher level，the genetic base seems to be much more for an all purpose learning rather than the more specific responses of imprinting．Chimpanzees，for instance，generally 【14】very good mother but Jane Goodall reports that some chimps carry the infant upside down or【15】fail to nurture the young．She believes that these females were the youngest or the 【16】child of a mother．In such circumstances，they did not have the opportunity to observe how their own mother【17】for her young．Certainly adolescent chimps who are still with their mothers when other young are born take much interest in the rearing of their young brother or sister．They have an excellent opportunity to learn，and the social ties that are created between mother and young 【18】 Goodall to describe the social unit as a family，The mother offspring tie is beyond 【19】；there is some evidence to【20】 that ties also continue between siblings of the same sex，that is “brother-brother” and “sister-sister”，
【9】
选项 A、appropriated
B、substituted
C、assigned
D、distributed
答案B
解析 考查惯用搭配。substitute A for B，意思是“用A代替B”，符合句意，故substituted为正确答案。
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