He must have heard of the bad news, ______can be perceived in his expression.

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问题 He must have heard of the bad news, ______can be perceived in his expression.

选项 A、as
B、what
C、while
D、that

答案A

解析 在这里作关系代词,引导非限定性定语从句,说明前面整句话,符合题意,故选A。
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